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American depository receipts of European stocks were 0.4% lower at 139.18 on the Bank of New York Mellon Europe ADR Index on Thursday morning.

Decliners in Continental Europe were led by Zealand Pharma ( ZEAL ), 8.5% lower, followed by Cellectis ( CLLS ), a French genome engineering company, down by 8.3% and Oasmia Pharmaceutical ( OASM ), a Sweden-based pharmaceutical company, 3.8% lower. Grifols ( GRFS ), a manufacturer of biopharmaceuticals, was down by 2.9%.

In the UK, Randgold Resources ( GOLD ),a gold mining company, was 6% lower, telecommunications major BT Group (BT) was down by 4.1% and Nightstar (NITE), a clinical-stage gene therapy company, was 3.8% lower. Lloyds Banking Group (LYG) was down by 3%.

Gainers in Continental Europe were led by Trivago (TRVG), an online hotel search platform operator, 8.5% higher, followed by Credit Suisse (CS), a Swiss financial services company, up by 5.2% and Ascendis Pharma (ASND), a Danish clinical stage biopharmaceutical company, 4.4% higher. Telecom Italia (TI), an Italian telecommunications company, was up by 2.9%.

And, in the UK, BHP Billiton (BBL), a mining company, was 1.2% higher, Carnival (CUK), a cruise vacation company, was up by 1.1% and Fly Leasing (FLY), an aircraft leasing company, was also 1.1% higher. Amarin (AMRN), a biopharmaceutical company, was up by 0.9%.

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